Herbs, perennial, with moderately to strongly thickened rootstock. Stems erect, 12–43 cm, uniformly strigose. Leaves opposite; stipules distinct, linear-subulate or nodiform to papilliform nodiform to papilliform stipules often reddish brown, 0.1–0.7 mm, strigose; petiole 0.3–1.2 mm, strigose; blade: proximal ovate to ovate-elliptic, distal linear to elliptic-linear, 7–41 × 2–5 mm, distal leaf blades more than 6 times as long as wide, base asymmetric, cuneate to rounded, margins entire, often involute on drying, apex acute, surfaces usually short strigose, occasionally glabrous adaxially; venation pinnate, only midvein conspicuous. Cyathia solitary at distal nodes; peduncle 1.1–2.4 mm. Involucre turbinate to campanulate-turbinate, 1–1.5 × 1–1.4 mm, strigose; glands 4, green to yellow-green, concave, narrowly oblong, 0.2–0.4 × 0.4–0.7 mm; appendages white, flabellate, 0.5–1.1 × 0.3–0.5 mm, distal margin shallowly and irregularly toothed. Staminate flowers 16–26. Pistillate flowers: ovary strigose; styles 0.3–0.5 mm, 2-fid at apex to almost 1/2 length. Capsules broadly ovoid, 2–2.6 × 2.5–3 mm, strigose; columella 1.6–2.2 mm. Seeds white, ovoid, 4-angled in cross section, 1.7–2.2 × 1.1–1.2 mm, transversely low-ridged or wrinkled.